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G.C. 2015 — Gamindustri has entered a precarious season known as the CPU Shift Period. During this time of social unrest, deprecating rumors about the Goddesses have begun to saturate Gamindustri. Neptune and the others worry the citizens will soon call for new leaders to replace them and end their rule.
In these critical moments for Planeptune, Neptune and Nepgear mysteriously disappear. Their destination: a divergent world, nearing its catastrophic end, called the Zero Dimension. They encounter the last remaining Goddess of this world, Uzume Tennouboshi, who is desperately waging a lonely battle against a colossal evil known as the Dark CPU.
Will Neptune and Nepgear be able to work together with Uzume in order to save Zero Dimension from tragedy? So begins a new adventure for Neptune…